Let's see if I can answer all of the questions:
Happens on different lenses on all three bodies. Doesn't appear to play favorites that way with lenses or combinations.
Exposure mode = AV and TAV that I've noticed so far. Can try others.
In-camera mode from two in a sequence:
https://emmisoure.zenfolio.com/p104379158 (I paid for zenfolio but have yet to build anything there - ugh!) - think I've set it to be downloadable (original .jpgs)
Often occurs when panning or tracking birds/insects. Sometimes, though, when I think I'm holding still, too. Burst shots of flowers, birds, etc.
Could be a sticky blade problem. But unlikely. I've swapped out lenses among bodies. And it seems to be a body rather than a lens problem. I'll explore that further, though. (to MetteHHH)
Re 4x Kx bodies - I hate swapping lenses. And my wife and I both shoot. So I try to collect the current body that I'm shooting. And then refrain from swapping lenses so much. Mostly I shoot bees, birds, flowers, and gardens. Nothing too special about any of that.
Shutter count of one is 100k+. The other two are...I think...~20K or so. (16K on the posted .jpg. - the other is roughly similar)
I'm 99pct sure it's not an issue with lenses. I've used so many (some new) among the cameras and it does seem body specific. In fact, after the bees on the muscari, I put a lens that's had blown-out results onto the "good" body and got consistently-lit shots. I can fiddle further there.
Thanks for the replies so far. Let me know if I've missed an answer or have not been clear.