Originally posted by leekil
Based on Occam's razor, I would think that the way it "knows where the film starts" is that there is a standard length of paper ahead of the film; it always advances the same amount each time, and doesn't have any sort of sensor to detect where the film is.
---------- Post added 10-07-17 at 10:25 AM ----------
The fact that you line up the Start arrow when you load the film seems to support my assumption.
I have both the 645N and 645N II and have used the Pentax 645 in the past, and this is exactly how they work. The manual and mechanical Rolleiflex has the function the OP mentioned where it senses the change in thickness when the film proper starts, but most other medium format film cameras "knows here the film starts" based on the lenght of the leader.